The Foundations of the Origin of Species. Darwin Charles

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СКАЧАТЬ that domestication produces little effect for several generations in rendering the organization plastic; but afterwards, as if by an accumulated effect, the original character of the species suddenly gives way or breaks.

      On Selection

      We have hitherto only referred to the first appearance in individuals of new peculiarities; but to make a race or breed, something more is generally198 requisite than such peculiarities (except in the case of the peculiarities being the direct effect of constantly surrounding conditions) should be inheritable, – namely the principle of selection, implying separation. Even in the rare instances of sports, with the hereditary tendency very strongly implanted, crossing must be prevented with other breeds, or if not prevented the best characterized of the half-bred offspring must be carefully selected. Where the external conditions are constantly tending to give some character, a race possessing this character will be formed with far greater ease by selecting and breeding together the individuals most affected. In the case of the endless slight variations produced by the indirect effects of domestication on the action of the reproductive system, selection is indispensable to form races; and when carefully applied, wonderfully numerous and diverse races can be formed. Selection, though so simple in theory, is and has been important to a degree which can hardly be overrated. It requires extreme skill, the results of long practice, in detecting the slightest difference in the forms of animals, and it implies some distinct object in view; with these requisites and patience, the breeder has simply to watch for every the smallest approach to the desired end, to select such individuals and pair them with the most suitable forms, and so continue with succeeding generations. In most cases careful selection and the prevention of accidental crosses will be necessary for several generations, for in new breeds there is a strong tendency to vary and especially to revert to ancestral forms: but in every succeeding generation less care will be requisite for the breed will become truer; until ultimately only an occasional individual will require to be separated or destroyed. Horticulturalists in raising seeds regularly practise this, and call it “roguing,” or destroying the “rogues” or false varieties. There is another and less efficient means of selection amongst animals: namely repeatedly procuring males with some desirable qualities, and allowing them and their offspring to breed freely together; and this in the course of time will affect the whole lot. These principles of selection have been methodically followed for scarcely a century; but their high importance is shown by the practical results, and is admitted in the writings of the most celebrated agriculturalists and horticulturalists; – I need only name Anderson, Marshall, Bakewell, Coke, Western, Sebright and Knight.

      Even in well-established breeds the individuals of which to an unpractised eye would appear absolutely similar, which would give, it might have been thought, no scope to selection, the whole appearance of the animal has been changed in a few years (as in the case of Lord Western’s sheep), so that practised agriculturalists could scarcely credit that a change had not been effected by a cross with other breeds. Breeders both of plants and animals frequently give their means of selection greater scope, by crossing different breeds and selecting the offspring; but we shall have to recur to this subject again.

      The external conditions will doubtless influence and modify the results of the most careful selection; it has been found impossible to prevent certain breeds of cattle from degenerating on mountain pastures; it would probably be impossible to keep the plumage of the wild-duck in the domesticated race; in certain soils, no care has been sufficient to raise cauliflower seed true to its character; and so in many other cases. But with patience it is wonderful what man has effected. He has selected and therefore in one sense made one breed of horses to race and another to pull; he has made sheep with fleeces good for carpets and other sheep good for broadcloth; he has, in the same sense, made one dog to find game and give him notice when found, and another dog to fetch him the game when killed; he has made by selection the fat to lie mixed with the meat in one breed and in another to accumulate in the bowels for the tallow-chandler199; he has made the legs of one breed of pigeons long, and the beak of another so short, that it can hardly feed itself; he has previously determined how the feathers on a bird’s body shall be coloured, and how the petals of many flowers shall be streaked or fringed, and has given prizes for complete success; – by selection, he has made the leaves of one variety and the flower-buds of another variety of the cabbage good to eat, at different seasons of the year; and thus has he acted on endless varieties. I do not wish to affirm that the long-and short-wooled sheep, or that the pointer and retriever, or that the cabbage and cauliflower have certainly descended from one and the same aboriginal wild stock; if they have not so descended, though it lessens what man has effected, a large result must be left unquestioned.

      In saying as I have done that man makes a breed, let it not be confounded with saying that man makes the individuals, which are given by nature with certain desirable qualities; man only adds together and makes a permanent gift of nature’s bounties. In several cases, indeed, for instance in the “Ancon” sheep, valuable from not getting over fences, and in the turnspit dog, man has probably only prevented crossing; but in many cases we positively know that he has gone on selecting, and taking advantage of successive small variations.

      Selection200 has been methodically followed, as I have said, for barely a century; but it cannot be doubted that occasionally it has been practised from the remotest ages, in those animals completely under the dominion of man. In the earliest chapters of the Bible there are rules given for influencing the colours of breeds, and black and white sheep are spoken of as separated. In the time of Pliny the barbarians of Europe and Asia endeavoured by cross-breeding with a wild stock to improve the races of their dogs and horses. The savages of Guyana now do so with their dogs: such care shows at least that the characters of individual animals were attended to. In the rudest times of English history, there were laws to prevent the exportation of fine animals of established breeds, and in the case of horses, in Henry VIII’s time, laws for the destruction of all horses under a certain size. In one of the oldest numbers of the Phil. Transactions, there are rules for selecting and improving the breeds of sheep. Sir H. Bunbury, in 1660, has given rules for selecting the finest seedling plants, with as much precision as the best recent horticulturalist could. Even in the most savage and rude nations, in the wars and famines which so frequently occur, the most useful of their animals would be preserved: the value set upon animals by savages is shown by the inhabitants of Tierra del Fuego devouring their old women before their dogs, which as they asserted are useful in otter-hunting201: who can doubt but that in every case of famine and war, the best otter-hunters would be preserved, and therefore in fact selected for breeding. As the offspring so obviously take after their parents, and as we have seen that savages take pains in crossing their dogs and horses with wild stocks, we may even conclude as probable that they would sometimes pair the most useful of their animals and keep their offspring separate. As different races of men require and admire different qualities in their domesticated animals, each would thus slowly, though unconsciously, be selecting a different breed. As Pallas has remarked, who can doubt but that the ancient Russian would esteem and endeavour to preserve those sheep in his flocks which had the thickest coats. This kind of insensible selection by which new breeds are not selected and kept separate, but a peculiar character is slowly given to the whole mass of the breed, by often saving the life of animals with certain characteristics, we may feel nearly sure, from what we see has been done by the more direct method of separate selection within the last 50 years in England, would in the course of some thousand years produce a marked effect.

      Crossing Breeds

      When once two or more races are formed, or if more than one race, or species fertile inter se, originally existed in a wild state, their crossing becomes a most copious source of new races202. When two well-marked races are crossed the offspring in the first generation take more or less after either parent or are quite intermediate between them, or rarely assume characters in some degree new. In the second and several succeeding generations, СКАЧАТЬ