Origin of Cultivated Plants. Alphonse de Candolle

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СКАЧАТЬ great abundance of a species is no proof of its antiquity. Agave Americana, so common on the shores of the Mediterranean, although introduced from America, and our cardoon, which now covers a great part of the Pampas of La Plata, are remarkable instances in point. As a rule, an invading species makes rapid way, while extinction is, on the contrary, the result of the strife of several centuries against unfavourable circumstances.12

      The designation which should be adopted for allied species, or, to speak scientifically, allied forms, is a problem often presented in natural history, and more often in the category of cultivated species than in others. These plants are changed by cultivation. Man adopts new and convenient forms, and propagates them by artificial means, such as budding, grafting, the choice of seeds, etc. It is clear that, in order to discover the origin of one of these species, we must eliminate as far as possible the forms which appear to be artificial, and concentrate our attention on the others. A simple reflection may guide this choice, namely, that a cultivated species varies chiefly in those parts for which it is cultivated. The others remain unmodified, or present trifling alterations, of which the cultivator takes no note, because they are useless to him. We may expect, therefore, to find the fruit of a wild fruit tree small and of a doubtfully agreeable flavour, the grain of a cereal in its wild state small, the tubercles of a wild potato small, the leaves of indigenous tobacco narrow, etc., without, however, going so far as to imagine that the species developed rapidly under cultivation, for man would not have begun to cultivate it if it had not from the beginning presented some useful or agreeable qualities.

      When once a cultivated plant has been reduced to such a condition as permits of its being reasonably compared with analogous spontaneous forms, we have still to decide what group of nearly similar plants it is proper to designate as constituting a species. Botanists alone are competent to pronounce an opinion on this question, since they are accustomed to appreciate differences and resemblances, and know the confusion of certain works in the matter of nomenclature. This is not the place to discuss what may reasonably be termed a species. I have stated in some of my articles the principles which seem to me the best. As their application would often require a study which has not been made, I have thought it well occasionally to treat quasi-specific forms as a group which appears to me to correspond to a species, and I have sought the geographical origin of these forms as though they were really specific.

      To sum up: botany furnishes valuable means of guessing or proving the origin of cultivated plants and for avoiding mistakes. We must, however, by no means forget that practical observation must be supplemented by research in the study. After gaining information from the collector who sees the plants in a given spot or district, and who draws up a flora or a catalogue of species, it is indispensable to study the known or probable geographical distribution in books and in herbaria, and to reflect upon the principles of geographical botany and on the questions of classification, which cannot be done by travelling or collecting. Other researches, of which I shall speak presently, must be combined with those of botany if we would arrive at satisfactory conclusions.

      3. Archæology and Palæontology. The most direct proof which can be conceived of the ancient existence of a species in a given country is to see its recognizable fragments in old buildings or deposits, of a more or less certain date.

      The fruits, seeds, and different portions of plants taken from ancient Egyptian tombs, and the drawings which surround them in the pyramids, have given rise to most important researches, which I shall often have to mention. Nevertheless, there is a possible source of error; the fraudulent introduction of modern plants into the sarcophagi of the mummies. This was easily discovered in the case of some grains of maize, for instance, a plant of American origin, which were introduced by the Arabs; but species cultivated in Egypt within the last two or three thousand years may have been added, which would thus appear to have belonged to an earlier period. The tumuli or mounds of North America, and the monuments of the ancient Mexicans and Peruvians, have furnished records about the plants cultivated in that part of the world. Here we are concerned with an epoch subsequent to the pyramids of Egypt.

      The deposits of the Swiss lake-dwellings have been the subject of important treatises, among which that of Heer, quoted just now, holds the first place. Similar works have been published on the vegetable remains found in other lakes or peat mosses of Switzerland, Savoy, Germany, and Italy. I shall quote them with reference to several species. Dr. Gross has been kind enough to send me seeds and fruits taken from the lake-dwellings of Neuchâtel; and my colleague, Professor Heer, has favoured me with several facts collected at Zurich since the publication of his work. I have already said that the rubbish-heaps of the Scandinavian countries, called kitchen-middens, have furnished no trace of cultivated vegetables.

      The tufa of the south of France contains leaves and other remains of plants, which have been discovered by MM. Martins, Planchon, de Saporta, and other savants. Their date is not, perhaps, always earlier than that of the first lacustrine deposits, and it is possible that it agrees with that of ancient Egyptian monuments, and of ancient Chinese books. Lastly, the mineralogic strata, with which geologists are specially concerned, tell us much about the succession of vegetable forms in different countries; but here we are dealing with epochs far anterior to agriculture, and it would be a strange and certainly a most valuable chance if a modern cultivated species were discovered in the European tertiary epoch. No such discovery has hitherto been made with any certainty, though uncultivated species have been recognized in strata prior to the glacial epoch of the northern hemisphere. For the rest, if we do not succeed in finding them, the consequences will not be clear, since it may be said, either that such a plant came at a later date from a different region, or that it had formerly another form which renders its recognition impossible in a fossil state.

      4. History. Historical records are important in order to determine the date of certain cultures in each country. They also give indications as to the geographical origin of plants when they have been propagated by the migrations of ancient peoples, by travellers, or by military expeditions.

      The assertions of authors must not, however, be accepted without examination.

      The greater number of ancient historians have confused the fact of the cultivation of a species in a country with that of its previous existence there in a wild state. It has been commonly asserted, even in our own day, that a species cultivated in America or China is a native of America or China. A no less common error is the belief that a species comes originally from a given country because it has come to us from thence, and not direct from the place in which it is really indigenous. Thus the Greeks and Romans called the peach the Persian apple, because they had seen it cultivated in Persia, where it probably did not grow wild. It was a native of China, as I have elsewhere shown. They called the pomegranate, which had spread gradually from garden to garden from Persia to Mauritania, the apple of Carthage (Malum Punicum). Very ancient authors, such as Herodotus and Berosius, are yet more liable to error, in spite of their desire to be accurate.

      We shall see, when we speak of maize, that historical documents which are complete forgeries may deceive us about the origin of a species. It is curious, for it seems to be no one’s interest to lie about such agricultural facts. Fortunately, facts of botany and archæology enable us to detect errors of this nature.

      The principal difficulty, which commonly occurs in the case of ancient historians, is to find the exact translation of the names of plants, which in their books always bear the common names. I shall speak presently of the value of these names and how the science of language may be brought to bear on the questions with which we are occupied, but I must first indicate those historical notions which are most useful in the study of cultivated plants.

      Agriculture came originally, at least so far as the principal species are concerned, from three great regions, in which certain plants grew, regions which had no communication with each other. These are – China, the south-west of Asia (with Egypt), and intertropical America. I do not mean to say that in Europe, in Africa, and elsewhere savage tribes may not have cultivated a few species locally, СКАЧАТЬ