Mutual Aid: A Factor of Evolution. kniaz Petr Alekseevich Kropotkin

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СКАЧАТЬ of studying the animal world in the wide uninhabited regions of Northern Asia and East Russia; and it is impossible to study like regions without being brought to the same ideas. I recollect myself the impression produced upon me by the animal world of Siberia when I explored the Vitim regions in the company of so accomplished a zoologist as my friend Polyakoff was. We both were under the fresh impression of the Origin of Species, but we vainly looked for the keen competition between animals of the same species which the reading of Darwin's work had prepared us to expect, even after taking into account the remarks of the third chapter (p. 54). We saw plenty of adaptations for struggling, very often in common, against the adverse circumstances of climate, or against various enemies, and Polyakoff wrote many a good page upon the mutual dependency of carnivores, ruminants, and rodents in their geographical distribution; we witnessed numbers of facts of mutual support, especially during the migrations of birds and ruminants; but even in the Amur and Usuri regions, where animal life swarms in abundance, facts of real competition and struggle between higher animals of the same species came very seldom under my notice, though I eagerly searched for them. The same impression appears in the works of most Russian zoologists, and it probably explains why Kessler's ideas were so welcomed by the Russian Darwinists, whilst like ideas are not in vogue amidst the followers of Darwin in Western Europe.

      The first thing which strikes us as soon as we begin studying the struggle for existence under both its aspects—direct and metaphorical—is the abundance of facts of mutual aid, not only for rearing progeny, as recognized by most evolutionists, but also for the safety of the individual, and for providing it with the necessary food. With many large divisions of the animal kingdom mutual aid is the rule. Mutual aid is met with even amidst the lowest animals, and we must be prepared to learn some day, from the students of microscopical pond-life, facts of unconscious mutual support, even from the life of micro-organisms. Of course, our knowledge of the life of the invertebrates, save the termites, the ants, and the bees, is extremely limited; and yet, even as regards the lower animals, we may glean a few facts of well-ascertained cooperation. The numberless associations of locusts, vanessae, cicindelae, cicadae, and so on, are practically quite unexplored; but the very fact of their existence indicates that they must be composed on about the same principles as the temporary associations of ants or bees for purposes of migration. As to the beetles, we have quite well-observed facts of mutual help amidst the burying beetles (Necrophorus). They must have some decaying organic matter to lay their eggs in, and thus to provide their larvae with food; but that matter must not decay very rapidly. So they are wont to bury in the ground the corpses of all kinds of small animals which they occasionally find in their rambles. As a rule, they live an isolated life, but when one of them has discovered the corpse of a mouse or of a bird, which it hardly could manage to bury itself, it calls four, six, or ten other beetles to perform the operation with united efforts; if necessary, they transport the corpse to a suitable soft ground; and they bury it in a very considerate way, without quarrelling as to which of them will enjoy the privilege of laying its eggs in the buried corpse. And when Gleditsch attached a dead bird to a cross made out of two sticks, or suspended a toad to a stick planted in the soil, the little beetles would in the same friendly way combine their intelligences to overcome the artifice of Man. The same combination of efforts has been noticed among the dung-beetles.

      Even among animals standing at a somewhat lower stage of organization we may find like examples. Some land-crabs of the West Indies and North America combine in large swarms in order to travel to the sea and to deposit therein their spawn; and each such migration implies concert, co-operation, and mutual support. As to the big Molucca crab (Limulus), I was struck (in 1882, at the Brighton Aquarium) with the extent of mutual assistance which these clumsy animals are capable of bestowing upon a comrade in case of need. One of them had fallen upon its back in a corner of the tank, and its heavy saucepan-like carapace prevented it from returning to its natural position, the more so as there was in the corner an iron bar which rendered the task still more difficult. Its comrades came to the rescue, and for one hour's time I watched how they endeavoured to help their fellow-prisoner. They came two at once, pushed their friend from beneath, and after strenuous efforts succeeded in lifting it upright; but then the iron bar would prevent them from achieving the work of rescue, and the crab would again heavily fall upon its back. After many attempts, one of the helpers would go in the depth of the tank and bring two other crabs, which would begin with fresh forces the same pushing and lifting of their helpless comrade. We stayed in the Aquarium for more than two hours, and, when leaving, we again came to cast a glance upon the tank: the work of rescue still continued! Since I saw that, I cannot refuse credit to the observation quoted by Dr. Erasmus Darwin—namely, that "the common crab during the moulting season stations as sentinel an unmoulted or hard-shelled individual to prevent marine enemies from injuring moulted individuals in their unprotected state."(5)

      Facts illustrating mutual aid amidst the termites, the ants, and the bees are so well known to the general reader, especially through the works of Romanes, L. Buchner, and Sir John Lubbock, that I may limit my remarks to a very few hints.(6) If we take an ants' nest, we not only see that every description of work-rearing of progeny, foraging, building, rearing of aphides, and so on—is performed according to the principles of voluntary mutual aid; we must also recognize, with Forel, that the chief, the fundamental feature of the life of many species of ants is the fact and the obligation for every ant of sharing its food, already swallowed and partly digested, with every member of the community which may apply for it. Two ants belonging to two different species or to two hostile nests, when they occasionally meet together, will avoid each other. But two ants belonging to the same nest or to the same colony of nests will approach each other, exchange a few movements with the antennae, and "if one of them is hungry or thirsty, and especially if the other has its crop full … it immediately asks for food." The individual thus requested never refuses; it sets apart its mandibles, takes a proper position, and regurgitates a drop of transparent fluid which is licked up by the hungry ant. Regurgitating food for other ants is so prominent a feature in the life of ants (at liberty), and it so constantly recurs both for feeding hungry comrades and for feeding larvae, that Forel considers the digestive tube of the ants as consisting of two different parts, one of which, the posterior, is for the special use of the individual, and the other, the anterior part, is chiefly for the use of the community. If an ant which has its crop full has been selfish enough to refuse feeding a comrade, it will be treated as an enemy, or even worse. If the refusal has been made while its kinsfolk were fighting with some other species, they will fall back upon the greedy individual with greater vehemence than even upon the enemies themselves. And if an ant has not refused to feed another ant belonging to an enemy species, it will be treated by the kinsfolk of the latter as a friend. All this is confirmed by most accurate observation and decisive experiments.(7)

      In that immense division of the animal kingdom which embodies more than one thousand species, and is so numerous that the Brazilians pretend that Brazil belongs to the ants, not to men, competition amidst the members of the same nest, or the colony of nests, does not exist. However terrible the wars between different species, and whatever the atrocities committed at war-time, mutual aid within the community, self-devotion grown into a habit, and very often self-sacrifice for the common welfare, are the rule. The ants and termites have renounced the "Hobbesian war," and they are the better for it. Their wonderful nests, their buildings, superior in relative size to those of man; their paved roads and overground vaulted galleries; their spacious halls and granaries; their corn-fields, harvesting and "malting" of grain;(8) their, rational methods of nursing their eggs and larvae, and of building special nests for rearing the aphides whom Linnaeus so picturesquely described as "the cows of the ants"; and, finally, their courage, pluck, and, superior intelligence—all these are the natural outcome of the mutual aid which they practise at every stage of their busy and laborious lives. That mode of life also necessarily resulted in the development of another essential feature of the life of ants: the immense development of individual initiative which, in its turn, evidently led to the development of that high and varied intelligence which cannot but strike the human observer.(9)

      If we knew no other facts from animal life than what we know about the ants and the termites, we already might safely conclude that mutual aid (which leads to mutual confidence, the first condition for courage) and individual initiative (the first СКАЧАТЬ