Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras. Alfred Russel Wallace

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СКАЧАТЬ than one, or in certain exceptional areas where deep ocean currents carry the débris of land to greater distances.^[18]

      If we now examine the stratified rocks found in the very centre of all our great continents, we find them to consist of sandstones, limestones, conglomerates, or shales, which must, as we have seen, have been deposited within a comparatively short distance of a sea-shore. Sir Archibald Geikie says:—"Among the thickest masses of sedimentary rock—those of the ancient Palæozoic systems—no features recur more continually than the alternations of different sediments, and the recurrence of surfaces covered with well-preserved ripple-marks, trails and burrows of annelides, polygonal and irregular desiccation marks, like the cracks at the bottom of a sun-dried muddy pool. These phenomena unequivocally point to shallow and even littoral waters. They occur from bottom to top of formations, which reach a thickness of several thousand feet. They can be interpreted only in one way, viz., that the formations in question began to be laid down in shallow water; that during their formation the area of deposit gradually subsided for thousands of feet; yet that the rate of accumulation of sediment kept pace on the whole with this depression; and hence that the original shallow-water character of the deposits remained, even after the original sea-bottom had been buried under a vast mass of sedimentary matter." He goes on to say, that this general statement applies to the more recent as well as to the more ancient formations, and concludes—"In short, the more attentively the stratified rocks of the earth are studied, the more striking becomes the absence of any formations among them, which can legitimately be considered those of a deep sea. They have all been deposited in comparatively shallow water."^[19]

      The arrangement and succession of the stratified rocks also indicate the mode and place of their formation. We find them stretching across the country in one general direction, in belts of no great width though often of immense length, just as we should expect in shore deposits; and they often thin out and change from coarse to fine in a definite manner, indicating the position of the adjacent land from the débris of which they were originally formed. Again quoting Sir Archibald Geikie:—"The materials carried down to the sea would arrange themselves then as they do still, the coarser portions nearest the shore, the finer silt and mud furthest from it. From the earliest geological times the great area of deposit has been, as it still is, the marginal belt of sea-floor skirting the land. It is there that nature has always strewn the dust of continents to be."

      The Movements of Continents.—As we find these stratified rocks of different periods spread over almost the whole surface of existing continents where not occupied by igneous or metamorphic rocks, it follows that at one period or another each part of the continent has been under the sea, but at the same time not far from the shore. Geologists now recognise two kinds of movements by which the deposits so formed have been elevated into dry land—in the one case the strata remain almost level and undisturbed, in the other they are contorted and crumpled, often to an enormous extent. The former often prevails in plains and plateaus, while the latter is almost always found in the great mountain ranges. We are thus led to picture the land of the globe as a flexible area in a state of slow but incessant change; the changes consisting of low undulations which creep over the surface so as to elevate and depress limited portions in succession without perceptibly affecting their nearly horizontal position; and also of intense lateral compression, supposed to be produced by partial subsidence along certain lines of weakness in the earth's crust, the effect of which is to crumple the strata and force up certain areas in great contorted masses, which, when carved out by subaërial denudation into peaks and valleys, constitute our great mountain systems.^[20] In this way every part of a continent may again and again have sunk beneath the sea, and yet as a whole may never have ceased to exist as a continent or a vast continental archipelago. And, as subsidence will always be accompanied by deposition, of sediments from the adjacent land, piles of marine strata many thousand feet thick may have been formed in a sea which was never very deep, by means of a slow depression either continuous or intermittent, or through alternate subsidences and elevations, each of moderate amount.

      Supposed Oceanic Formations;—the Origin of Chalk.—There seems very good reason to believe that few, if any, of the rocks known to geologists correspond exactly to the deposits now forming at the bottom of our great oceans. The white oceanic mud, or Globigerina-ooze, found in all the great oceans at depths varying from 250 to nearly 3,000 fathoms, and almost constantly in depths under 2,000 fathoms, has, however, been supposed to be an exception, and to correspond exactly to our white and grey chalk. Hence some naturalists have maintained that there has probably been one continuous formation of chalk in the Atlantic from the Cretaceous epoch to the present day. This view has been adopted chiefly on account of the similarity of the minute organisms found to compose a considerable proportion of both deposits, more especially the pelagic Foraminifera, of which several species of Globigerina appear to be identical in the chalk and the modern Atlantic mud. Other extremely minute organisms whose nature is doubtful, called coccoliths and discoliths, are also found in both formations, while there is a considerable general resemblance between the higher forms of life. Sir Wyville Thomson tells us, that—"Sponges are abundant in both, and the recent chalk-mud has yielded a large number of examples of the group porifera vitrea, which find their nearest representatives among the Ventriculites of the white chalk. The echinoderm fauna of the deeper parts of the Atlantic basin is very characteristic, and yields an assemblage of forms which represent in a remarkable degree the corresponding group in the white chalk. Species of the genus Cidaris are numerous; some remarkable flexible forms of the Diademidæ seem to approach Echinothuria."^[21] Now, as some explanation of the origin of chalk had long been desired by geologists, it is not surprising that the amount of resemblance shown to exist between it and some kinds of oceanic mud should have been at once seized upon, and the conclusion arrived at that chalk is a deep-sea oceanic formation exactly analogous to that which has been shown to cover large areas of the Atlantic, Pacific and Southern oceans.

      But there are several objections to this view which seem fatal to its acceptance. In the first place, no specimens of Globigerina-ooze from the deep ocean-bed yet examined agree even approximately with chalk in chemical composition, only containing from 44 to 79 per cent. of carbonate of lime, with from 5 to 11 per cent of silica, and from 8 to 33 per cent. of alumina and oxide of iron.^[22] Chalk, on the other hand, contains usually from 94 to 99 per cent. of carbonate of lime, and a very minute quantity of alumina and silica. This large proportion of carbonate of lime implies some other source of this mineral, and it is probably to be found in the excessively fine mud produced by the decomposition and denudation of coral reefs. Mr. Dana, the geologist of the United States Exploring Expedition, found in the elevated coral reef of Oahu, one of the Sandwich Islands, a deposit closely resembling chalk in colour, texture, &c.; while in several growing reefs a similar formation of modern chalk undistinguishable from the ancient, was observed.^[23] Sir Charles Lyell well remarks that the pure calcareous mud produced by the decomposition of the shelly coverings of mollusca and zoophytes would be much lighter than argillaceous or arenaceous mud, and being thus transported to greater distances would be completely separated from all impurities.

      Now the Globigerinæ have been shown by the Challenger explorations to abound in all moderately warm seas; living both at the surface, at various depths in the water, and at the bottom. It was long thought that they were surface-dwellers only, and that their dead tests sank to the bottom, producing the Globigerina-ooze in those areas where other deposits were absent or scanty. But the examination of the whole of the dredgings and surface-gatherings of the Challenger by Mr. H. B. Brady has led him to a different conclusion; for he finds numerous forms at the bottom quite distinct from those which inhabit the surface, while, when the same species live both at surface and bottom, the latter are always larger and have thicker and stronger cell-walls. This view is also supported by the fact that in many stations not far from our own shores Globigerinæ are abundant in bottom dredgings, but are never found on the surface in the towing-nets.^[24] These organisms then exist almost universally where the waters are pure and are not too cold, and they would naturally abound most where the diffusion of carbonate of lime both in suspension and solution afforded them an abundant supply of material for their shelly coverings. Dr. Wallich believes СКАЧАТЬ