The Natural Philosophy of Love. Remy de Gourmont

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СКАЧАТЬ where it disappears in the mollusk series, whereof some possess so luxurious a love-organism. The simple and very naive form, that in which the sperm and the eggs are produced simultaneously inside the same individual, is found only in inferior organisms. Normal parthenogenesis belong equally to summary and to complicated animals, to wheel-animals and to bees. Among arthropodes, that is to say among insects in general, the sexes are always separate, save in certain tardigrade arachnids, but these are the ones which offer the finest cases of parthenogenesis, generation without aid of the male. The term need not be taken literally. For as there is no indefinite scissiparity without coupling, there is no unlimited parthenogenesis without fecundation: the female is fecundated for several generations which transmit this power, but there comes a day when the female who has not encountered a male gives birth to males and females. They couple and produce females parthenogenetically endowed. This has been for long time a mystery—it is still a mystery, for side by side with normal parthenogenesis there is irregular parthenogenesis, there are cases where non-fecundated eggs behave exactly as fecundated eggs, without anyone's knowing why.

      The virgin-begotten cycle of plant-lice is famous, that of wheel-animals not less entertaining. The males, smaller than the females, live but two or three days, couple and die. The fecundated females lay eggs whence come nothing but females, unless the eggs are subjected to a temperature above 18 degrees (centigrade); above that the eggs hatch out males. Between the periods of coupling there are long stretches of virgin-birth, nothing but females producing females, until the temperature permits a male hatch. In two years the plant louse runs through ten or twelve parthenogeneses; in July of the second year, there appear winged individuals, these are still female, but double size, and they lay two sets of eggs, whereof the smaller hatch male (the male is three or four times smaller than the female), the larger eggs hatch female; there is coupling and the cycle begins again.

      For long people believed the plant louse truly androgynous. Réaumur and Bonnet, having seen isolated plant-lice reproduce themselves were convinced of this, when Trembley, a man of genius, celebrated also for his observations of hydra, threw out the idea: Who knows whether a coupling of these lice does not fecundate them for several generations? He had discovered the basis of parthenogenesis. Facts upheld him. Bonnet described the male and female, and noted even the genital ardour of this sticking leaf-louse, this milch-cow of the ants.

      Parthenogenesis is a sign-post. Nothing more clearly demonstrates the importance of the male or the precision of his function. The female appears to be the whole show, without the male she is nothing. She is the machine and has to be wound up to go. The male is merely the key. People have tried to obtain fecundation by false keys. Eggs of sea-anemones, and star-fish have been hatched by contact with exciting chemicals, acids, alkalines, sugar, salt, alcohol, ether, chloroform, strychnine gas, carbonic acid. But one has never been able to bring these scientific larvæ to maturity, and everything leads one to believe that if one succeeded, and that if these artificial beings were capable of reproduction, it would be but for a limited period. This provoked parthenogenesis is neither more nor less interesting than the normal. It is doubtless abnormal, but abnormal parthenogenesis is not infrequent in nature; eggs of the bombyx, of star-fish, and of frogs, hatch sometimes without fecundation, and very probably because they have accidentally come up against the very stimulant which the excellent experimenters have lavished on them. Whether sperm acts as an "excitant" or as fecundator, the action is no easier to understand by one label than by another. The queen bee lays both fecundated and non-fecundated eggs; the first hatch female, the second invariably male, here the male element would seem to be the product of parthenogenesis and the female to require previous fecundation. In contrast, among plant-lice, the generations of female continue for nearly two years. There is an order in these things, as in all things, but it is not yet apparent; one notes only, that however long and varied be the parthenogenetic period, it is limited somewhere by the necessity of the female principle being united with the male principle. After all, hereditary fecundation is no more extraordinary than particular fecundation, it is a mode of perpetuating life which the exercise of one's reason should make one consider as perfectly normal.

      One ought, at the end of this summary chapter, to be courageous enough to say that fecundation, as vulgarly understood, is merely an illusion. Taking man and woman (or no matter what dioic metazoaire) the man does not fecundate the woman; what happens is at once more mysterious and more simple. From the male A, the great Male, and from the great Female B are born without any fecundation whatever, spontaneously, little males a and little females 6. These little males are called spermatozoides, and the little females, ovules; it is between these new creatures, between these spores, that the fecundating union occurs. One then observes that a and b resolve themselves into a third animal x, which by natural growth becomes either A or B. Then the cycle begins again. The union between A and B is merely a preparation; A and B are nothing but channels carrying a and b, carrying them often far beyond themselves. Like the plant-lice or drones, the mammifers called man are subject to alternate generation, one parthenogenesis always separating the veritable conjunction of the differentiated elements. Coupling is not fecundation; it is merely the mechanism; its utility is merely in that it puts two parthenogenetic products into relation. This relation occurs inside the female, or outside the female (as in case of fishes); the milieu has an importance of fact, not of principle.

      CHAPTER IV

      SEXUAL DIMORPHISM

       Table of Contents

      I. Invertebrates: formation of the male.—Primitivity of the female.—Minuscule males: the bonellie.—Regression of the male into the male organ: the cirripedes.—Generality of sexual dimorphism.—Superiority of the female in most insect species.—Exceptions.—Numeric dimorphism.—Female hymenoptera.—Multiplicity of her activities.—Male's purely sexual rôle.—Dimorphism of ants and termites.—Grasshoppers and crickets.—Spiders.—Coleoptera.—Glow-worm.—Cochineal's strange dimorphism.

      I. Invertebrates.—At a moment fairly undecided in the general evolution the male organ specializes into the male individual. Religious symbolisms may or might have been intended to mean this. The female is primitive. At the third month, the human embryo has external uro-genital organs clearly resembling the female organs. To arrive at complete female estate they need undergo but a very slight modification; to become male they have to undergo a considerable and very complex transformation. The external genital organs of the female are not, as has been often said, the product of an arrested development; quite the contrary, the male organs undergo a supplementary development, which is moreover useless, for the penis is a luxury and a danger: the bird who does without it is no less wanton thereby.

      One finds general proof of the female's primitivity in the extreme smallness of certain male invertebrates, so tiny indeed that one can only consider them as autonomous masculine organs, or even as spermatozoides. The male of the syngames (an internal parasite of birds) is less a creature than an appendix; he remains in constant contact with the organs of the female, stuck obliquely into her side, and justifying the name "two-headed worm" which has been given to this wretched and duplex animalculus. The female bonellie is a sea worm shaped like a sort of cornucopia sack fifteen centimetres in length: the male is represented by a minuscule filament of about one or two millimetres, that is to say about one-thousandth her size. Each female supports about twenty. These males live, first in the œsophagus, then descend into the oviduct where they impregnate the eggs. Only their very definite function clears them from the charge of being parasites; in fact they were long supposed to be parasites, while men sought vainly for the male of the prodigious bonellie.

      Side by side with males who are merely individualized sexual organs, one sees males who have lost nearly all organs save the male organ itself. Certain hermaphrodite cirripedes (mollusks attached by a peduncle [stalk]) cling as parasites to the coat of other cirripedes: whence a diminution of volume, a regression of ovaries, abolition of nutritive functions; the stalk takes root in the living, nourishing СКАЧАТЬ