Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras. Alfred Russel Wallace

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СКАЧАТЬ the gorgeous macaws; the curassows, the trumpeters, and the sun-bitterns. Here again there is no other continent or region that can produce such an assemblage of remarkable and perfectly distinct groups of birds; and no less wonderful is its richness in species, since these fully equal, if they do not surpass, those of the two great tropical regions of the Eastern Hemisphere (the Ethiopian and the Oriental) combined.

      As an additional indication of the distinctness and isolation of the Neotropical region from all others, and especially from the whole Eastern Hemisphere, we must say something of the otherwise widely distributed groups which are absent. Among mammalia we have first the order Insectivora, entirely absent from South America, though a few species are found in Central America and the West Indies; the Viverridæ or civet family is wholly wanting, as are every form of sheep, oxen, or antelopes; while the swine, the elephants, and the rhinoceroses of the old world are represented by the diminutive peccaries and tapirs.

      Among birds we have to notice the absence of tits, true flycatchers, shrikes, sunbirds, starlings, larks (except a solitary species in the Andes), rollers, bee-eaters, and pheasants, while warblers are very scarce, and the almost cosmopolitan wagtails are represented by a single species of pipit.

      We must also notice the preponderance of low or archaic types among the animals of South America. Edentates, marsupials, and rodents form the majority of the terrestrial mammalia; while such higher groups as the carnivora and hoofed animals are exceedingly deficient. Among birds a low type of Passeres, characterised by the absence of the singing muscles, is excessively prevalent, the enormous groups of the ant-thrushes, tyrants, tree-creepers, manakins, and chatterers belonging to it. The Picariæ (a lower group) also prevail to a far greater extent than in any other regions, both in variety of forms and number of species; and the chief representatives of the gallinaceous birds—the curassows and tinamous, are believed to be allied, the former to the brush-turkeys of Australia, the latter (very remotely) to the ostriches, two of the least developed types of birds.

      Whether, therefore, we consider its richness in peculiar forms of animal life, its enormous variety of species, its numerous deficiencies as compared with other parts of the world, or the prevalence of a low type of organisation among its higher animals, the Neotropical region stands out as undoubtedly the most remarkable of the great zoological divisions of the earth.

      In reptiles, amphibia, fresh-water fishes, and insects, this region is equally peculiar, but we need not refer to these here, our only object now being to establish by a sufficient number of well-known and easily remembered examples, the distinctness of each region from all others, and its unity as a whole. The former has now been sufficiently demonstrated, but it may be well to say a few words as to the latter point.

      The only outlying portions of the region about which there can be any doubt are—Central America, or that part of the region north of the Isthmus of Panama, the Antilles or West Indian Islands, and the temperate portion of South America including Chili and Patagonia.

      In Central America, and especially in Mexico, we have an intermixture of South American and North American animals, but the former undoubtedly predominate, and a large proportion of the peculiar Neotropical groups extend as far as Costa Rica. Even in Guatemala and Mexico we have howling and spider-monkeys, coati-mundis, tapirs, and armadillos; while chatterers, manakins, ant-thrushes, and other peculiarly Neotropical groups of birds are abundant. There is therefore no doubt as to Mexico forming part of this region, although it is comparatively poor, and exhibits the intermingling of temperate and tropical forms.

      The West Indies are less clearly Neotropical, their poverty in mammals as well as in most other groups being extreme, while great numbers of North American birds migrate there in winter. The resident birds, however, comprise trogons, sugar-birds, chatterers, with many humming-birds and parrots, representing eighteen peculiar Neotropical genera; a fact which decides the region to which the islands belong.

South temperate America is also very poor as compared with the tropical parts of the region, and its insects contain a considerable proportion of north temperate forms. But it contains armadillos, cavies and opossums; and its birds all belong to American groups, though, owing to the inferior climate and deficiency of forests, a number of the families of birds peculiar to tropical America are wanting. Thus there are no manakins, chatterers, toucans, trogons, or motmots; but there are abundance of hang-nests, tyrant-birds, ant-thrushes, tree-creepers, and a fair proportion of humming-birds, tanagers and parrots. The zoology is therefore thoroughly Neotropical, although somewhat poor; and it has a number of peculiar forms of strictly Neotropical types—as the chinchillas, alpacas, &c., which are not found in the tropical regions except in the high Andes.

      Comparison of Zoological Regions with the Geographical Divisions of the Globe.—Having now completed our survey of the great zoological regions of the globe, we find that they do not differ so much from the old geographical divisions as our first example might have led us to suppose. Europe, Asia, Africa, Australia, North America, and South America, really correspond, each to a zoological region, but their boundaries require to be modified more or less considerably; and if we remember this, and keep their extensions or limitations always in our mind, we may use the terms "South American" or "North American," as being equivalent to Neotropical and Nearctic, without much inconvenience, while "African" and "Australian" equally well serve to express the zoological type of the Ethiopian and Australian regions. Europe and Asia require more important modifications. The European fauna does indeed well represent the Palæarctic in all its main features, and if instead of Asia we say tropical Asia we have the Oriental region very fairly defined; so that the relation of the geographical with the zoological primary divisions of the earth is sufficiently clear. In order to make these relations visible to the eye and more easily remembered, we will put them into a tabular form:

Regions.	Geographical Equivalent.
Palæarctic	Europe, with north temperate Africa and Asia.
Ethiopian	Africa (south of the Sahara) with Madagascar.
Oriental	Tropical Asia, to Philippines and Java.
Australian	Australia, with Pacific Islands, Moluccas, &c.
Nearctic	North America, to North Mexico.
Neotropical	South America, with tropical N. America and W. Indies.

      The following arrangement of the regions will indicate their geographical position, and to a considerable extent their relation to each other.

      N e a r c t i c--P a l æ a r c t i c | | |

       | | Oriental | Ethiopian |

       Neo- |

       Tropical Australian

CHAPTER IV

       Table of Contents

      EVOLUTION THE KEY TO DISTRIBUTION

      Importance of the Doctrine of Evolution—The Origin of New Species—Variation in Animals—The Amount of Variation in North American Birds—How New Species arise from a Variable Species—Definition and Origin of Genera—Cause of the Extinction of Species—The Rise and Decay of Species and Genera—Discontinuous Specific Areas, why Rare—Discontinuity of the Area of Parus Palustris—Discontinuity of Emberiza Schœniclus—The European and Japanese Jays—Supposed СКАЧАТЬ