The Principles of Biology, Volume 1 (of 2). Spencer Herbert

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СКАЧАТЬ truth, rendered so familiar by infinite repetition that we forget its significance, must be named. There is invariably, and necessarily, a conformity between the vital functions of any organism and the conditions in which it is placed – between the processes going on inside of it and the processes going on outside of it. We know that a fish cannot live long in air, or a man under water. An oak growing in the ocean and a seaweed on the top of a hill, are incredible combinations of ideas. We find that each kind of animal is limited to a certain range of climate; each kind of plant to certain zones of latitude and elevation. Of the marine flora and fauna, each species is found only between such and such depths. Some blind creatures flourish in dark caves; the limpet where it is alternately covered and uncovered by the tide; the red-snow alga rarely elsewhere than in the arctic regions or among alpine peaks.

      Grouping together the cases first named, in which a particular change in the circumstances of an organism is followed by a particular change in it, and the cases last named, in which the constant actions occurring within an organism imply some constant actions occurring without it; we see that in both, the changes or processes displayed by a living body are specially related to the changes or processes in its environment. And here we have the needful supplement to our conception of Life. Adding this all-important characteristic, our conception of Life becomes – The definite combination of heterogeneous changes, both simultaneous and successive, in correspondence with external co-existences and sequences. That the full significance of this addition may be seen, it will be necessary to glance at the correspondence under some of its leading aspects.17

      § 28. Neglecting minor requirements, the actions going on in a plant pre-suppose a surrounding medium containing at least carbonic acid and water, together with a due supply of light and a certain temperature. Within the leaves carbon is being appropriated and oxygen given off; without them, is the gas from which the carbon is taken, and the imponderable agents that aid the abstraction. Be the nature of the process what it may, it is clear that there are external elements prone to undergo special re-arrangements under special conditions. It is clear that the plant in sunshine presents these conditions and so effects these re-arrangements. And thus it is clear that the changes which primarily constitute the plant's life, are in correspondence with co-existences in its environment.

      If, again, we ask respecting the lowest protozoon how it lives; the answer is, that while on the one hand its substance is undergoing disintegration, it is on the other hand absorbing nutriment; and that it may continue to exist, the one process must keep pace with, or exceed, the other. If further we ask under what circumstances these combined changes are possible, there is the reply that the medium in which the protozoon is placed, must contain oxygen and food – oxygen in such quantity as to produce some disintegration; food in such quantity as to permit that disintegration to be made good. In other words – the two antagonistic processes taking place internally, imply the presence externally of materials having affinities that can give rise to them.

      Leaving those lowest animal forms which simply take in through their surfaces the nutriment and oxygenated fluids coming in contact with them, we pass to those somewhat higher forms which have their tissues slightly specialized. In these we see a correspondence between certain actions in the digestive sac, and the properties of certain surrounding bodies. That a creature of this order may continue to live, it is necessary not only that there be masses of substance in the environment capable of transformation into its own tissue, but also that the introduction of these masses into its stomach, shall be followed by the secretion of a solvent fluid which will reduce them to a fit state for absorption. Special outer properties must be met by special inner properties.

      When, from the process by which food is digested, we turn to the process by which it is seized, the same general truth faces us. The stinging and contractile power of a polype's tentacle, correspond to the sensitiveness and strength of the creatures serving it for prey. Unless that external change which brings one of these creatures in contact with the tentacle, were quickly followed by those internal changes which result in the coiling and drawing up of the tentacle, the polype would die of inanition. The fundamental processes of integration and disintegration within it, would get out of correspondence with the agencies and processes without it, and the life would cease.

      Similarly, when the creature becomes so large that its tissue cannot be efficiently supplied with nutriment by mere absorption through its lining membrane, or duly oxygenated by contact with the fluid bathing its surface, there arises a need for a distributing system by which nutriment and oxygen may be carried throughout the mass; and the functions of this system, being subsidiary to the two primary functions, form links in the correspondence between internal and external actions. The like is obviously true of all those subordinate functions, secretory and excretory, that facilitate oxidation and assimilation.

      Ascending from visceral actions to muscular and nervous actions, we find the correspondence displayed in a manner still more obvious. Every act of locomotion implies the expenditure of certain internal forces, adapted in amounts and directions to balance or out-balance certain external forces. The recognition of an object is impossible without a harmony between the changes constituting perception, and particular properties co-existing in the environment. Escape from enemies implies motions within the organism, related in kind and rapidity to motions without it. Destruction of prey requires a special combination of subjective actions, fitted in degree and succession to overcome a group of objective ones. And so with those countless automatic processes constituting instincts.

      In the highest order of vital changes the same fact is equally manifest. The empirical generalization that guides the farmer in his rotation of crops, serves to bring his actions into concord with certain of the actions going on in plants and soil. The rational deductions of the educated navigator who calculates his position at sea, form a series of mental acts by which his proceedings are conformed to surrounding circumstances. Alike in the simplest inferences of the child and the most complex ones of the man of science, we find a correspondence between simultaneous and successive changes in the organism, and co-existences and sequences in its environment.

      § 29. This general formula which thus includes the lowest vegetal processes along with the highest manifestations of human intelligence, will perhaps call forth some criticisms which it is desirable here to meet.

      It may be thought that there are still a few inorganic actions included in the definition; as, for example, that displayed by the mis-named storm-glass. The feathery crystallization which, on a certain change of temperature, takes place in its contained solution, and which afterwards dissolves to reappear in new forms under new conditions, may be held to present simultaneous and successive changes that are to some extent heterogeneous, that occur with some definiteness of combination, and, above all, occur in apparent correspondence with external changes. In this case vegetal life is simulated to a considerable extent; but it is merely simulated. The relation between the phenomena occurring in the storm-glass and in the atmosphere respectively, is not a correspondence at all, in the proper sense of the word. Outside there is a thermal change; inside there is a change of atomic arrangement. Outside there is another thermal change; inside there is another change of atomic arrangement. But subtle as is the dependence of each internal upon each external change, the connexion between them does not, in the abstract, differ from the connexion between the motion of a straw and the motion of the wind that disturbs it. In either case a change produces a change, and there it ends. The alteration wrought by some environing agency on this or any other inanimate object, does not tend to induce in it a secondary alteration which anticipates some secondary alteration in the environment. But in every living body there is a tendency towards secondary alterations of this nature; and it is in their production that the correspondence consists. The difference may be best expressed by symbols. Let A be a change in the environment, and B some resulting change in an inorganic mass. Then A having produced B, the action ceases. Though the change A in the environment is followed by some consequent change a in it; no parallel sequence in the inorganic mass simultaneously generates in it some change b that has reference to the change a. But if we take a living body of the requisite organization, and let the change A impress СКАЧАТЬ